Europe’s most complete stegosaurian skull unearthed in Teruel, Spain

A new study by palaeontologist from the Fundación Dinópolis, published in the journal Vertebrate Zoology, rewrites the evolutionary history of this dino group.

A photo of a person holding a large, textured fossil resembling a dinosaur skull, with distinct eye sockets, against a backdrop of grass.
Close-up photograph of the Dacentrurus armatus skull found in Riodeva (Teruel, Spain).

Palaeontologists from the Fundación Conjunto Paleontológico de Teruel-Dinópolis have published new research in the prestigious scientific journal Vertebrate Zoology published by Senckenberg. The article describes a partial stegosaurian skull discovered in the municipality of Riodeva (Teruel, Spain) and proposes a new hypothesis about the evolutionary history of plated dinosaurs.

Stegosaurs were dinosaurs mainly characterized by being plant-eaters, moving on all fours, and displaying two rows of plates and/or spines from the neck to the end of the tail. The specimen studied was recovered during the palaeontological excavations led by the Fundación Dinópolis at the “Están de Colón” fossil site, located in sediments of the Villar del Arzobispo Formation, dating to the Late Jurassic epoch (around 150 million years ago). It is the best-preserved stegosaurian skull ever found in Europe and has been identified as belonging to the species Dacentrurus armatus.

An illustration of a  large, spiked dinosaur with a long neck standing by the shore, roaring against a backdrop of sea and cloudy sky.
Illustration of Dacentrurus armatus. Credit: Adrián Blázquez / Fundación Dinópolis.

Sergio Sánchez Fenollosa, a researcher at the Fundación Dinópolis and co-author of the study, explains: “The detailed study of this exceptional fossil has allowed us to reveal previously unknown aspects of the anatomy of Dacentrurus armatus, the quintessential European stegosaur, which in 2025 marks 150 years since its first description. Dinosaurian skulls are rarely preserved due to the extreme fragility of their bones. This discovery is key to understanding how stegosaurian skulls evolved.

A close-up photo of a dinosaur sculpture's head, with another dinosaur in the background.
Head of a Dacentrurus armatus sculpture on display at Dinópolis (Teruel, Spain).

Furthermore, alongside the anatomical study, we have also proposed a new hypothesis that redefines the evolutionary relationships of stegosaurs worldwide. As a result of this work, we have formalized the definition of a new group called Neostegosauria”.

According to the researchers, this new group includes medium to large-sized stegosaurian species that at least lived in what is now Africa and Europe during the Middle and Late Jurassic, in North America during the Late Jurassic, and in Asia during the Late Jurassic and Early Cretaceous.

Alberto Cobos, managing director of the Fundación Dinópolis and co-author of the publication, adds: “This dual achievement–both the study of an exceptional fossil and the proposal of a new evolutionary hypothesis–positions this research as a global reference in stegosaurian studies.

This fossil site from Riodeva continues to be a subject of research and still holds numerous relevant fossils, including more postcranial elements from the same adult specimen and, notably, juvenile individuals, a particularly rare combination in this type of dinosaurs. These discoveries continue to exponentially increase the palaeontological heritage of the province of Teruel, making it one of the iconic regions for understanding the evolution of life on Earth”.

3D rendering of a fossilized piece of dinosaur skull.
Cranial reconstruction of a stegosaur, showing the studied fossil in its anatomical position.

The scientific article has been published in the renowned international journal Vertebrate Zoology under the title New insights into the phylogeny and skull evolution of stegosaurian dinosaurs: An extraordinary cranium from the European Late Jurassic (Dinosauria: Stegosauria). The authors are palaeontologists Sergio Sánchez Fenollosa and Alberto Cobos from the Fundación Dinópolis. The article is available open access at: https://doi.org/10.3897/vz.75.e146618

Illustration of a Dacentrurus skull from Riodeva, Spain, showing three views: fossil, outlined diagram, and 3D model, scale 10 cm.
Set of images of the most complete stegosaurian skull found in Europe, belonging to Dacentrurus armatus and recovered from Riodeva (Teruel, Spain).

This contribution from the Fundación Dinópolis, affiliated with the Dept. of Medio Ambiente y Turismo of the Gobierno de Aragón, is part of the activities of Research Group E04-23R FOCONTUR, funded by the Gobierno de Aragón (through the Dept. of Empleo, Ciencia y Universidades). Additionally, it is part of the research of the Unidad de Paleontología de Teruel, funded by the Gobierno de España (through the Ministry of Ciencia, Innovación y Universidades). The work at this site is also supported by the project titled Los yacimientos paleontológicos de la provincia de Teruel como factor de desarrollo territorial (IV), subsidized by the Gobierno de España and the Gobierno de Aragón through the Teruel Investment Fund (via the Dept. of Presidencia, Economía y Justicia).

Research article:

Sánchez-Fenollosa S, Cobos A (2025) New insights into the phylogeny and skull evolution of stegosaurian dinosaurs: An extraordinary cranium from the European Late Jurassic (Dinosauria: Stegosauria). Vertebrate Zoology 75: 165-189. https://doi.org/10.3897/vz.75.e146618

Image credit: Fundación Conjunto Paleontológico de Teruel-Dinópolis