Why two prehistoric sharks found in Ohio got new names

Research leads to rediscovery of forgotten fossils.

Until recently, Orthacanthus gracilis could have been considered the “John Smith” of prehistoric shark names, given how common it was.

Three different species of sharks from the late Paleozoic Era – about 310 million years ago – were mistakenly given that same name, causing lots of grief to paleontologists who studied and wrote about the sharks through the years and had trouble keeping them apart.

But now Loren Babcock, a professor of earth sciences at The Ohio State University, has finished the arduous task of renaming two of the three sharks – and in the process rediscovered a wealth of fossil fishes that had been stored at an Ohio State museum for years but had been largely forgotten.

Loren Babcock with a collection of Orton Museum’s fossil fishes, including several from John Newberry. Photo by The Ohio State University

In order to change the names, Babcock had to go through a process governed by the International Commission on Zoological Nomenclature (ICZN). He had to document the need to change the names, propose new names and submit them to an ICZN-recognized journal for peer review and then have the ICZN officially accept the names.

Tooth of the shark Orthacanthus lintonensis. The tooth is about 13 mm long.

“It was one of the most complex naming problems we have had in paleontology, which is probably one reason no one attempted to fix it until now,” Babcock said.

“A lot of scientists in the field have written, thanking me for doing this. We are all happy it is finally done,” he said.

One measure of the impact the renaming has had on the field: Babcock’s paper announcing the new names was just published in the journal ZooKeys on Jan. 8, but it has already been referenced on seven different Wikipedia pages.

The original Orthacanthus gracilis fossil was found in Germany and named in 1848. That species gets to keep the name.

The remaining two fossils were found in Ohio and named by the famous American paleontologist John Strong Newberry in 1857 and 1875.

Portrait of John Strong Newberry

Babcock renamed the Ohio sharks Orthacanthus lintonensis and Orthacanthus adamas, both based on the name of the place where they were originally found.

Why did Newberry give the two Ohio sharks the same name?

“He probably just forgot. It was nearly 20 years between the time the two species were named,” Babcock said.

And as far as giving it the same name as a German species: “In those days, it was really difficult to search for names that were already in existence – they did not have the internet.”

The sharks themselves were fascinating creatures, Babcock said.  They were large and creepy, nearly 10 feet long, and looked more like eels than present-day sharks, with long dorsal fins extending the length of their backs and a peculiar spine extending backward from their heads.

They lived in the fresh or brackish water of what are known as “coal swamps” of the late Carboniferous Period (323-299 million years ago) during the late Paleozoic Era. They belong to an extinct group of chondrichthyans (which includes sharks, skates and rays) called the xenacanthiforms.

Dorsal spine of Orthacanthus adamas. The spine is about 71 mm long.

Newberry was for a time the chief geologist at the Geological Survey of Ohio. He played an important role in the early growth of what is now the Orton Geological Museum at Ohio State.

Babcock, who is the current director of the Orton Museum, decided to begin the renaming process after reviewing the museum’s collection. He was surprised to see how many fossils the museum had that had been collected by Newberry, including the two prehistoric sharks.

Babcock wrote about Orton’s Newberry collection in a new article published in the Journal of Vertebrate Paleontology.

Through the years, scientists have written about how various Newberry specimens had been lost. It turns out many had been at the Orton Museum.

“No museum has a larger collection of Newberry’s fossils except for the American Museum of Natural History in New York City,” Babcock said.

“Not a lot of people are aware of that – I did not even know the extent of our collection. If you’re looking for part of the Newberry collection and can’t find it in the American Museum of Natural History, it is probably going to be here.”

Research article:

Babcock LE (2024) Replacement names for two species of Orthacanthus Agassiz, 1843 (Chondrichthyes, Xenacanthiformes), and discussion of Giebelodus Whitley, 1940, replacement name for Chilodus Giebel, 1848 (Chondrichthyes, Xenacanthiformes), preoccupied by Chilodus Müller & Troschel, 1844 (Actinopterygii, Characiformes). ZooKeys 1188: 219-226. https://doi.org/10.3897/zookeys.1188.108571

News piece originally published by the Ohio State University. Republished with permission.

Follow ZooKeys on Facebook and X.

New species of 65-million-year-old shark ‘accidentally’ discovered in Alabama

The shark lived shortly after the extinction of the dinosaurs, and was discovered from a box of teeth collected over 100 years ago.

A new species of shark, which lived shortly after the mass extinction of the dinosaurs, was discovered when palaeontologist Jun Ebersole came across a 100-year-old box of teeth at the Geological Survey in Alabama, USA.

“Having documented hundreds of fossil fish species over the last decade, I found it puzzling that these teeth were from a shark that I didn’t recognise,” says Ebersole, Director of Collections, McWane Science Center, Birmingham, AL, who quickly realised the teeth belong to a new species.

New shark species tooth.
Palaeohypotodus bizzocoi tooth. Credit: McWane Science Center.

The team, consisting of Ebersole, David Cicimurri, Curator of Natural History, South Carolina State Museum in Columbia, and T. Lynn Harrell Jr., Palaeontologist and Fossil Collections Curator at the Geological Survey of Alabama in Tuscaloosa, published their discovery in the open-access journal Fossil Record.

The shark is a new species of Palaeohypotodus (pronounced pale-ee-oh-hype-oh-toe-duss), which means “ancient small-eared tooth,” in reference to the small needle-like fangs present on the sides of the teeth. Scientists believe it may have looked like a modern sand tiger shark.

Moder sand tiger shark.
Modern sand tiger. Credit: Wikimedia commons.

Living approximately 65-million-years ago in the Paleocene era, Palaeohypotodus bizzocoi was likely a leading predator as the oceans recovered following the death of the dinosaurs, when more than 75% of life on Earth went extinct.

In Alabama, much of the southern half of the state was covered by a shallow tropical to sub-tropical ocean during the Paleocene.

New shark species teeth.
Palaeohypotodus bizzocoi teeth. Credit: Ebersole et al.

“This time period is understudied, which makes the discovery of this new shark species that much more significant,” Harrell says. “Shark discoveries like this one give us tremendous insights into how ocean life recovers after major extinction events and also allows us to potentially forecast how global events, like climate change, affect marine life today.”

As part of their study of this ancient fish, the team compared the fossil teeth to those of various living sharks, like Great Whites and Makos. According to Cicimurri, shark teeth differ in shape depending on where they are located in the mouth.

“By studying the jaws and teeth of living sharks, it allowed us to reconstruct the dentition of this ancient species and showed that it had a tooth arrangement that differed from any living shark,” Cicimurri says.

Dr. Bruce Bizzoco.
The late Dr. Bruce Bizzoco (1949-2022), for whom the new species is named.
Credit: McWane Science Center.

The new species has been named Palaeohypotodus bizzocoi for the late Dr. Bruce Bizzoco (1949-2022) of Birmingham, AL. Bizzoco served as a Dean at Shelton State Community College, archaeologist, and was a long-time volunteer at McWane Science Center.

This discovery is part of an ongoing project led by Ebersole and Cicimurri to document Alabama’s fossil fishes. Together, they have confirmed over 400 unique species of fossil sharks and bony fishes, which, according to Ebersole, makes Alabama one of the richest places in the world in terms of fossil fish diversity.

Research paper

Ebersole JA, Cicimurri DJ, Harrell Jr. TL (2024) A new species of Palaeohypotodus Glickman, 1964 (Chondrichthyes, Lamniformes) from the lower Paleocene (Danian) Porters Creek Formation, Wilcox County, Alabama, USA. Fossil Record 27(1): 111-134. https://doi.org/10.3897/fr.27.e112800

Follow Fossil Record on X and Facebook.

Oldest family of jewel wasps discovered from Cretaceous amber in Lebanon

The new family, Protoitidae, and 10 new species are described in a new paper published in Journal of Hymenoptera Research.

Jewel wasps (Chalcidoidea) are one of the most diverse groups of insects, with more than 120,000 species described and an estimated true diversity of nearly one million. The chalcids are parasitoid wasps, which attack other insects to lay their eggs upon in order to feed and grow within the host. The evolution and origin of this immense group has puzzled scientists for decades. Based on fossils and molecular work, the group is thought to have originated in the late Jurassic around 162 million years ago. Until recently, no confirmed fossils were known from earlier than 100 million years ago. Now, a newly described family, Protoidae, provides the first glimpse into how these wasps appeared at the earliest stages of their evolution.

Protoita noyesi.

Scientists Jonah M. Ulmer, Dr. Petr Janšta, and Prof. Dr. Lars Krogmann, from SMNS – State Museum of Natural History Stuttgart, alongside Dr. Dany Azar from the Lebanese University describe the new family and 10 new species of jewel wasps in a paper in the open-access Journal of Hymenoptera Research.

The discovery of the family came about when one of the coauthors, Prof. Dr. Lars Krogmann, noticed an unusual fossil during a visit to the Natural History Museum in Paris. The specimen embedded in the amber had a long tail-like structure covering its ovipositor. “It was previously described as a completely different type of wasp, however the authors were quick to recognize it was indeed an ancient chalcid. Despite the prevalence of Chalcidoidea in the fossil record, none had ever been recorded from Lebanon or were known to be that old, nearly 130 million years old to be exact,” says Jonah Ulmer.

Cretaxenomerus brevis.

The researchers soon realized they had a new, and very old, family: currently the oldest known within the jewel wasps. “Multiple similar specimens in amber soon became apparent and the family now contains two genera, Protoita and Cretaxenomerus. The family name is derived from being a ‘proto’ form of the Chalcidoidea,” Ulmer explains.

These ancient species present a unique snapshot of what these wasps looked like in their earliest forms, this allows researchers to better understand the order of evolutionary events through time and when certain structures evolved that ultimately led to the massive diversity we see today.

Cretaxenomerus curvus.

The new family’s most striking characteristic is the long, shovel-like process, which extends from the end of the abdomen. While no living species have such a pronounced structure, it is hypothesized to have assisted with egg-laying and ovipositing or perhaps sifting through loose leaf litter for hosts.

The authors note that there are likely other equally old families of Chalcidoidea still lying in wait to be discovered, either in the ground or in old forgotten museum cabinets. “Protoitidae shows that we can keep looking further back in time than we expected and still find new, and old, species” says Ulmer.

Original source:

UImer JM, Janšta P, Azar D, Krogmann L (2023) At the dawn of megadiversity – Protoitidae, a new family of Chalcidoidea (Hymenoptera) from Lower Cretaceous Lebanese amber. Journal of Hymenoptera Research 96: 879-924. https://doi.org/10.3897/jhr.96.105494

Follow the Journal of Hymenoptera Research on Facebook and X.

Europe’s very own dinosaurs – the enigmatic Late Cretaceous rhabdodontids

Despite being widespread and abundant, these gregarious herbivores vanished in Western Europe around 69 million years ago due to environmental changes, while surviving longer in Eastern Europe.

When you think of dinosaurs, you might automatically imagine iconic dinosaurs as Tyrannosaurus and Triceratops. But at the same time when these were stomping on the ancient coastal plains of North America, some of their very distant cousins were reigning over Europe’s lands.

Life reconstructions and size comparison of three rhabdodontids. From left to right: Mochlodon suessi from eastern Austria (the smallest member of the group), Rhabdodon priscus from southern France (the largest member of the group), and Transylvanosaurus platycephalus from western Romania (the most recently named member of the group). Also shown is the silhoutte of a human (1.8 m tall) for scale. Reconstruction by Peter Nickolaus

During the Late Cretaceous (between 100 and 66 million years ago), Europe was an extensive archipelago with numerous small and large islands situated in a shallow tropical sea, the so-called Late Cretaceous European Archipelago. The dinosaur groups that lived on these islands were very different from those of other continents, often being much smaller than their mainland relatives. These European dinosaurs include small and medium-sized carnivorous theropods, armoured ankylosaurs, long-necked sauropods, duck-billed hadrosaurs, and rhabdodontids.

Skeletal reconstruction of Zalmoxes robustus. Scale bar: 20 cm

Arguably one of the most important of these European dinosaur groups is the family Rhabdodontidae, which groups together the most common medium-sized herbivores of the Late Cretaceous European Archipelago. A joint research team from the Universities of Tübingen (Germany), Budapest (Hungary) and Bucharest (Romania) recently reviewed what we know about these peculiar dinosaurs in a new paper published in the journal Fossil Record.

Generally, rhabdodontid dinosaurs were small to medium-sized animals with an overall body length of approximately 2–6 m. “They were probably habitually bipedal herbivores, characterised by a rather stocky build, with strong hind limbs, short forelimbs, a long tail, and a comparatively large, triangular skull that tapers anteriorly and ends in a narrow snout,” explains Felix Augustin, lead author of the study in Fossil Record.

“They had a relatively robust skull with strong jaws, large teeth and a pointy beak that was covered in keratin, demonstrating that these dinosaurs were well-adapted to eating tough plants.”

In some instances, fossil remains of several individuals of different ages have been found together, indicating that they were gregarious.

Although they died out well before the mass extinction in Western Europe (about 69 million years ago), potentially due to environmental changes that affected the plants they fed on, they survived much longer in Eastern Europe and were among the last non-avian dinosaurs still present before the end of the Cretaceous (66 million years ago).

Interestingly, fossils of rhabdodontids have only been found in Europe and only in rocks ranging in age from 86–66 million years ago, so they were endemic to the Late Cretaceous European Archipelago.

The group currently comprises nine different species from five European countries (France, Spain, Austria, Hungary, and Romania).

Palaeogeographic map of Europe during the latest Cretaceous (late Campanian), with the location of the most important rhabdodontid-bearing assemblages. 1 Transylvania, western Romania. 2 Iharkút, western Hungary. 3 Muthmannsdorf, eastern Austria. 4 Eastern southern France. 5 Western southern France. 6 Northern Spain. 7 Central Spain.

“The first rhabdodontid species was scientifically named more than 150 years ago and the last one as recently as November 2022, so, although the group looks back to a long research history, we still have much to learn about it,” says Felix Augustin.

“Generally, our portraying of the world of dinosaurs is heavily biased towards the well-known North-American and Asian dinosaur faunas,” he adds.

Type specimens of some rhabdodontid species. A. The original drawing of the lectotype of Rhabdodon priscus, MPLM 30, a partial left dentary. The specimen has since deteriorated (Pincemaille-Quillevere 2002). Modified after Matheron (1869). B. Holotype of Rhabdodon septimanicus, MDE D-30, an incomplete right dentary. Photo kindly provided by Eric Buffetaut. C. Lectotype of Mochlodon suessi, PIUW 2349/2, a right dentary. D. Holotype of Mochlodon vorosi, MTM V 2010.105.1, a left dentary. E. Holotype of Zalmoxes robustus, NHMUK R.3392, a right dentary. Photo kindly provided by János Magyar. F. Holotype right dentary of Zalmoxes shqiperorum, NHMUK R.4900. Note that the holotype of Z. shqiperorum also comprises several postcranial elements that presumably belong to the same individual as the dentary. Photo kindly provided by János Magyar.

Dinosaur fossils from the Late Cretaceous are much rarer in Europe than in North America or Asia, and thus far no complete skeleton of a rhabdodontid has been described. Even though they were so abundant and common in the Upper Cretaceous of Europe, several key aspects about them remain poorly known, including their detailed body proportions, their posture and locomotion, as well as their feeding behaviour.

“In the past decades, a wealth of new, and often well-preserved, rhabdodontid fossils has been discovered throughout Europe, the majority of which still remains to be studied,” says Felix Augustin. “A joint research project is currently underway to study the available fossil material in order to gain new insights into the evolution and lifestyle of these fascinating yet still poorly known dinosaurs.”

Original source:

Augustin FJ, Ősi A, Csiki-Sava Z (2023) The Rhabdodontidae (Dinosauria, Ornithischia), an enigmatic dinosaur group endemic to the Late Cretaceous European Archipelago. Fossil Record 26(2): 171-189. https://doi.org/10.3897/fr.26.108967

Follow Fossil Record on Facebook and Twitter:

Miniature Snail in a Rock Sandwich

First fossil record of thorn snails in the southern United States

Researchers from the USA and Switzerland, including Senckenberg scientist and first author Dr. Adrienne Jochum, have described the first fossil Carychium land snails from Florida. The rock layer containing the snail fossils, which are only a few millimeters in size, was accidentally uncovered during construction work and dates from the Pleistocene period between 2.58 million and 11,700 years ago. In their study, published in the open-access journal ZooKeys, the scientists also describe a previously unknown carychiid fossil species.
Light microscopic images of Carychium nashuaense.

The tiny snails of the genus Carychium with a maximum height of 2.5 millimeters and a width of 1.5 millimeters are known mostly east of the Mississippi River in the USA and from eastern Canada, Mexico and Jamaica as well as in sheltered humid habitats in Central America to Panama. “However, unlike the extant species, fossilized thorn snails are rarely found east of the Mississippi River. In our recent field work we have now provided the first fossil record of the genus in the southeastern United States, as well as the first fossil record ever for the species Carychium floridanum,” explains Dr. Adrienne Jochum of the Senckenberg Research Institute and Natural History Museum in Frankfurt and the Natural History Museum in Bern.

Fossil Carychium floridanum.

During the construction of a track bed for the Brightline railroad, which will connect Port Canaveral with Orlando International Airport, civil engineers accidentally came across a one-meter-thick layer of fossilized non-marine gastropods between two marine shell beds. “This ‘rock sandwich’ was formed during the Pleistocene, a geologic era characterized by repeated glaciations, climate changes, and fluctuations in water level that greatly influenced and shaped the region around present-day Florida. The shell layer is sandwiched between rock layers from the Lower Pleistocene, 2.58 to 0.77 million years ago, and the Upper Pleistocene, 140,000 to 120,000 years ago, and contains 14 freshwater and 28 terrestrial snail species.”

Later stage of excavation showing middle stratum of freshwater marl (c. 1 m thick) wedged between two layers of marine shell layers (each c. 3 m thick).

Among them is the snail Carychium floridanum, whose current representatives still live in humid, forested, and undisturbed habitats in central and northern Florida. The researchers also described a new species, Carychium nashuaense, which is less than 1.6 millimeters long and was previously unknown to science.

“To dislodge the fossil miniature snails from the rock layers, we first washed them through a graduated series of sieves. Next, 32 Carychium shells were culled under a microscope from a mixture of other mollusks and rock debris. A high-resolution X-ray tomograph helped us examine the spindle structure inside the fragile fossil shells and compare them with 3D reconstructions of the inner shell of still-living thorn snail species from the southeastern U.S., Mexico, Central America, and Jamaica,” explains Jochum.

While the design of the inner shell structure of Carychium floridanum has changed little from the Pleistocene to the present, the shell structure of Carychium nashuaense suggests a relationship with Central American Carychium relatives. “We suspect that the spread of the snails occurred via birds, mammals, and reptiles, who transported the small snails in their guts, fur, or feathers to the wetlands from which the alluvial sediments in the rock layer we studied originated. The subsequent mixing with other members of the genus led to the emergence of new species,” adds Jochum in conclusion.

***

Publication:

Jochum A, Bochud E, Haberthür D, Lee HG, Hlushchuk R, Portell RW (2023) Fossil Carychiidae (Eupulmonata, Ellobioidea) from the Lower Pleistocene Nashua Formation of Florida, with the description of a new species. ZooKeys 1167: 89-107.
https://doi.org/10.3897/zookeys.1167.102840

Press release originally published by Senckenberg. Republished with permission.

Follow ZooKeys on social media:

Oldest Pterodactylus fossil found in Germany

The fossil, just described in a study in the journal Fossil Record, is about one million years older than other Pterodactylus specimens.

Pterosaurs, the flying reptiles of the dinosaur era, originated in the Late Triassic (227 million years ago) and became extinct at the end-Cretaceous extinction event (66 million years ago). With wing spans ranging from 1 to 12 meters, they dominated the world’s skies for more than 160 million years.

The first described and named pterosaur – and namesake of the whole group – is Pterodactylus from the famous Solnhofen Limestone of Bavaria, southern Germany. Originally described in 1784 by the Italian naturalist Cosimo Alessandro Collini, the fossil was considered to be an aquatic animal for 25 years, before Georges Cuvier found out it was a flying reptile belonging to a new, previously unrecognized group.

The oldest specimen of this iconic pterosaur was recently found near Painten, a small town in the southern part of the Franconian Alb in central Bavaria. The fossil, described in a study in the journal Fossil Record, is about one million years older than other Pterodactylus specimens.

The specimen was unearthed in 2014 during excavations in an active limestone quarry. It took more than 120 hours of meticulous mechanical work using pneumatic tools and needles before the researchers could study it. The research team behind the discovery are Felix Augustin, Andreas Matzke, Panagiotis Kampouridis and Josephina Hartung from the University of Tübingen (Germany) and Raimund Albersdörfer from the Dinosaurier Museum Altmühltal (Germany).

“The rocks of the quarry, which yielded the new Pterodactylus specimen, consist of silicified limestone that has been dated to the upper Kimmeridgian stage (around 152 million years ago)”, explains Felix Augustin of the University of Tübingen, who is the lead author of the study. “Previously, Pterodactylus had only been found in younger rocks of southern Germany belonging to the Tithonian stage that follows after the Kimmeridgian”.

The specimen is a complete, well-preserved skeleton of a small-sized individual. “Only a very small portion of the left mandible as well as of the left and right tibia is missing. Otherwise, the skeleton is nearly perfectly preserved with every bone present and in its roughly correct anatomical position”, the researchers write in their study.

With a 5-cm-long skull, the Painten Pterodactylus represents a rare “sub-adult” individual. “Generally, the Pterodactylus specimens are not evenly distributed across the full size range but predominantly fall into distinct size-classes that are separated by marked gaps. The specimen from Painten is a rare representative of the first gap between the small and large sizes,” explains Augustin. “The Painten Pterodactylus was of an intermediate, and rarely found, ontogenetic age at the time of its death, between two consecutive year-classes.”

The Painten quarry has yielded many other “exquisitely preserved fossils”, including ichthyosaurs, turtles, marine and terrestrial crocodile-relatives, and dinosaurs.  Many of them, like this new pterosaur specimen, are on display in the new Dinosaurier Museum Altmühltal in Denkendorf (Bavaria, Germany).

Research article:

Augustin FJ, Kampouridis P, Hartung J, Albersdörfer R, Matzke AT (2022) The geologically oldest specimen of Pterodactylus: a new exquisitely preserved skeleton from the Upper Jurassic (Kimmeridgian) Plattenkalk deposits of Painten (Bavaria, Germany). Fossil Record 25(2): 331-343. https://doi.org/10.3897/fr.25.90692

Follow Fossil Record on Facebook and Twitter.

Jurassic Park in Eastern Morocco: Paleontology of the Kem Kem Group

The Kem Kem beds in Morocco are famous for the spectacular fossils found there, including at least four large-bodied non-avian theropods, several large-bodied pterosaurs and crocodilians. In their study, published in the open-access journal Zookeys, an international group of scientists, led by Dr. Nizar Ibrahim and Prof. Paul Sereno, evaluate the geological and paleontological significance of the study area.

The Kem Kem beds in Morocco are famous for the spectacular fossils found there, including at least four large-bodied non-avian theropods, several large-bodied pterosaurs and crocodilians.

Now, in a new geology and paleontology monograph, that reveals much more about the famous Kem Kem beds in Morocco, Dr. Nizar Ibrahim from the University of Detroit Mercy, Prof. Paul Sereno from the University of Chicago, and a team of international scholars from the United States, Europe and Morocco, have put together a comprehensive story that is published in the open-access journal Zookeys.

The aim of the new research is to provide the international community with an in-depth review of the rocks and fossils of the region. It reviews the geology and paleontology of this famous but surprisingly understudied area, describing and formally naming the strata and summarizing all of the preserved life forms, from fragile plants and insects to massive dinosaurs. The monograph also paints a picture of life as it once was some 95 million years ago by describing the paleoenvironments of the region, and the unusual predator-dominated fauna.

In 1996 Prof. Sereno and colleagues introduced the informal term “Kem Kem beds” for this fossil-rich escarpment. In this monograph, the authors review the original tri-level proposal for the region by French geologist Choubert (his informal “trilogie mésocretacée”) and propose the Kem Kem Group for the entire package of rock with two new names for the dinosaur-bearing layers based on the richest fossil sites, the Gara Sbaa and Douira formations.

The region is famous for the prodigious fossils found in all of these units, many derived from commercial fossil collecting, which obscures the precise location and level of the specimens. The monograph is the first work to pinpoint where many of the most important finds were made. Over the last 25 years in particular, paleontologists have brought to light a diverse array of new vertebrate fossils including at least four large-bodied non-avian theropods, several large-bodied pterosaurs, crocodilians, turtles and an array of sharks and bony fish.  

To put a comprehensive story together on the Kem Kem, the authors of the monograph visited collections of Kem Kem fossils around the world and led many expeditions to the region. Fossil and geological data reviewed in the monograph is derived from a number of different sources. A University of Chicago-led major expedition in 1995 generated a wealth of geological and paleontological data, as did later expeditions involving teams from the University College Dublin, the University of Portsmouth, the Faculté des Sciences Aïn Chock, the Muséum national d’Histoire naturelle, the University Cadi Ayyad, the Museo Civico di Storia Naturale (Milan), and the University of Detroit Mercy.

One of the key features of the Kem Kem assemblage is the presence of several large-bodied theropods, a group of dinosaurs that includes all of the meat-eaters. Most famous among these from the Kem Kem include the sail-backed Spinosaurus and the sabre-toothed Carcharodontosaurus.

Most fossils in the Kem Kem region are discovered as isolated fragmentary pieces weathered from sandstones. Only four partial dinosaur skeletons or skulls have been recovered, which include the long-necked sauropod Rebbachisaurus garasbae and the theropods Deltadromeus agilis, Carcharodontosaurus saharicus and Spinosaurus aegyptiacus. These Kem Kem theropods are among the largest known dinosaurian predators on record reaching adult body lengths in excess of 12 meters. 

“Given the continued input of new specimens and the continuing expansion of paleontological research, we predict that diversity in the Kem Kem Group will increase substantially in the coming decades. Based on our review of existing collections, this increase will include scores of taxa from the pond locality Oum Tkout including nonvertebrates, such as plants, insects, and ostracods, as well as an array of actinopterygian fish. We also anticipate a continuing trickle of new terrestrial vertebrates that will be named on better preserved specimens that are diagnostic at present only at the familial level, including turtles and various kinds of archosaurs. As nearly half of the reptilian families listed are indeterminate, better preserved specimens will offer future opportunities to recognize new reptilian genera” ,

share the authors.
Predators abound on land, in the air and in water some 95 million years on the shores of northern Africa —as shown by the abundant fossils in the Kem Kem region.
Large herbivores, such as the long-necked sauropod Rebbachisaurus, could have been hunted or scavenged by several large predators.
Credit: Artwork by Davide Bonadonna
License: CC-BY 4.0

“In summary, the Kem Kem assemblage of non-vertebrates and vertebrates is likely to continue to show dramatic increase in diversity in the coming decades. Nonetheless, the array of taxa currently known, which extends from plants across a range of aquatic and terrestrial vertebrates, is sufficiently mature to allow a summary of the vertebrate assemblage and a discussion of its paleoecological contex”,

conclude the researchers.

In his earlier research, a famous paleontologist from the University of Chicago Prof. Paul Sereno has described many outstanding dinosaur discoveries, including new Cretaceous crocodilians from the Sahara and two new fanged vegetarian dinosaur dwarfs (called heterodontosaurids). 


The most famous of Kem Kem dinosaurs, the semi-aquatic giant Spinosaurus, and the most common of Kem Kem fossils, the giant sawfish Onchopristis, tangle in the shallow coastal waters on a warm Late Cretaceous day.
Credit: Artwork by Davide Bonadonna
License: CC-BY 4.0

The wealth of aquatic life, including shrimp, bony fish, lungfish and giant lobe-finned coelacanths, supported a remarkable array of predators, including the fish-eating sail-backed Spinosaurus and toothless pterosaur Alanqa soaring overhead.
Credit: Artwork by Davide Bonadonna
License: CC-BY 4.0

Original source:

Ibrahim N, Sereno PC, Varricchio DJ, Martill DM, Dutheil DB, Unwin DM, Baidder L, Larsson HCE, Zouhri S, Kaoukaya A (2020) Geology and paleontology of the Upper Cretaceous Kem Kem Group of eastern Morocco. ZooKeys 928: 1-216. https://doi.org/10.3897/zookeys.928.47517

Contacts:

Prof. Paul Sereno
Email: dinosaur@uchicago.edu 

Dr. Nizar Ibrahim
Email: ibrahini@udmercy.edu

“Oldest bamboo” fossil from Eocene Patagonia turns out to be a conifer

A recent examination revealed that Chusquea oxyphylla, a fossilised leafy branch from the early Eocene in Patagonia, which has been widely cited as the oldest bamboo fossil and as evidence for a Gondwanan origin of bamboos is actually a conifer. The results of the finding are published in the open-access journal Phytokeys.

A fossilised leafy branch from the early Eocene in Patagonia described in 1941 is still often cited as the oldest bamboo fossil and the main fossil evidence for a Gondwanan origin of bamboos. However, a recent examination by Dr. Peter Wilf from Pennsylvania State University revealed the real nature of Chusquea oxyphylla. The recent findings, published in the paper in the open-access journal Phytokeys, show that it is actually a conifer.  

The corrected identification is significant because the fossil in question was the only bamboo macrofossil still considered from the ancient southern supercontinent of Gondwana. The oldest microfossil evidence for bamboo in the Northern Hemisphere belongs to the Middle Eocene, while other South American fossils are not older than Pliocene. 

Over the last decades, some authors have doubted whether the Patagonian fossil was really a bamboo or even a grass species at all. But despite its general significance, modern-day re-examinations of the original specimen were never published. Most scientists referring to it had a chance to study only a photograph found in the original publication from 1941 by the famous Argentine botanists Joaquín Frenguelli and Lorenzo Parodi.

In his recent study of the holotype specimen at Museo de La Plata, Argentina, Dr. Peter Wilf revealed that the fossil does not resemble members of the Chusquea genus or any other bamboo.

There is no evidence of bamboo-type nodes, sheaths or ligules. Areas that may resemble any bamboo features consist only of the broken departure points of leaf bases diverging from the twig. The decurrent, extensively clasping leaves are quite unlike the characteristically pseudopetiolate leaves of bamboos, and the heterofacially twisted free-leaf bases do not occur in any bamboo or grass,” wrote Dr. Wilf.

Instead, Wilf linked the holotype to the recently described fossils of the conifer genus Retrophyllum from the same fossil site, the prolific Laguna del Hunco fossil lake-beds in Chubut Province, Argentina. It matches precisely the distichous fossil foliage form of Retrophyllum spiralifolium, which was described based on a large set of data – a suite of 82 specimens collected from both Laguna del Hunco and the early middle Eocene Río Pichileufú site in Río Negro Province. 

Retrophyllum is a genus of six living species of rainforest conifers. Its habitat lies in both the Neotropics and the tropical West Pacific.

The gathered evidence firmly confirms that Chusquea oxyphylla has nothing in common with bamboos. Thus, it requires renaming.  Preserving the priority of the older name, Wilf combined Chusquea oxyphylla and Retrophyllum spiralifolium into Retrophyllum oxyphyllum.

The exclusion of a living New World bamboo genus from the overall floral list for Eocene Patagonia weakens the New World biogeographic signal of the late-Gondwanan vegetation of South America, which already showed much stronger links to living floras of the tropical West Pacific.

“The strongest New World signal remaining in Eocene Patagonia based on well-described macrofossils comes from fossil fruits of Physalis (a genus of flowering plants including tomatillos and ground cherries), which is an entirely American genus,” concludes Dr. Wilf.

Original source:
Wilf P (2020) Eocene “Chusquea” fossil from Patagonia is a conifer, not a bamboo. PhytoKeys 139: 77-89.
https://doi.org/10.3897/phytokeys.139.48717

The ‘Star dust’ wasp is a new extinct species named after David Bowie’s alter ego

During her study on fossil insects of the order Hymenoptera at China’s Capitol Normal University, student Longfeng Li visited the Smithsonian National Museum of Natural History, Washington, carrying two unidentified wasp specimens that were exceptionally well-preserved in Burmese amber. This type of fossilized tree resin is known for the quality of the fossil specimens which can be preserved inside it. Being 100 million years old, they provide an incredible view into the past.

The subsequent analysis of the specimens revealed that both represent species new to science. Furthermore, one of the wasps showed such amazing similarities to a modern group of wasps that it was placed in a currently existing genus, Archaeoteleiawhich has long been considered as an ancient lineage. The species are described in a study published in the open access Journal of Hymenoptera Research.

However, Archaeoteleia has changed since the times when the ancient wasp got stuck on fresh tree resin. The authors note that “a novice might not recognize the characters that unite the fossil with extant species”. For instance, the modern wasp species of the genus show visibly longer antennal segments and a different number of teeth on the mandible when compared to the fossil. In turn, the description of the new extinct species enhances the knowledge about living species by highlighting anatomical structures shared by all species within the genus.

This fossil wasp with living relatives received quite a curious name, Archaeoteleia astropulvis. The species name, astropulvis, translates from Latin to ‘star dust’. The discoverers chose the name to refer to both “the ancient source of the atoms that form our planet and its inhabitants”, as well as to commemorate the late David Bowie’s alter ego – Ziggy Stardust.

Unlike the Star dust wasp, the second new species belongs to a genus (Proteroscelio) known exclusively from Cretaceous fossils. Likewise, it is a tiny insect, measuring less than 2mm in length. It also plays an important role in taxonomy by expanding the anatomical diversity known from this extinct genus.

10388_Proteroscelio nexus

The authors conclude that their discovery, especially the Star dust wasp and its placement in an extant genus, where it is the only fossil species, “exemplifies the importance of understanding the extant fauna of a taxon to interpret fossils”.

“Such union of fossil and extant morphologies is especially illuminating and requires examination of both kinds of specimens,” they add.

###

Original source:

Talamas EJ, Johnson NF, Buffington ML, Dong R (2016) Archaeoteleia Masner in the Cretaceous and a new species of Proteroscelio Brues (Hymenoptera, Platygastroidea). In: Talamas EJ, Buffington ML (Eds) Advances in the Systematics of Platygastroidea. Journal of Hymenoptera Research 56: 241-261. https://doi.org/10.3897/jhr.56.10388