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Next-Generation Journal Publishing Platform ARPHA at Frankfurt Book Fair 2016

Following the launch of our self-developed journal publishing platform ARPHA (standing for Authoring, Reviewing, Publishing, Hosting and Archiving, all happening at one place), we were so happy with the outcome, that we couldn’t help sharing it with the world. Therefore, it’s on offer not only to our own journals and authors, but it’s also at hand to journals looking for their new home. Just let us know you’re interested!

On these lines, what could be a better place to have a chat about the transition in scholarly publishing, open science, research reproducibility and, of course, the advantages of having a journal published on ARPHA, than Frankfurt Book Fair 2016? Between 18th and 23rd October, this immense event will be all about unveiling and celebrating the evolution in the publishing industry, with exhibitors, trade and private visitors from across the globe, gathering together, led by their common expertise and passion.

Meet us at HotSpot Professional and Scientific Information Stage, Hall 4.2, Stand M90, during the Fair, and make sure you don’t miss the “ARPHA: Next-Generation Journal Publishing” presentation at 11:30 AM (local time) on Friday, 21st October, where Pensoft’s Founder and Managing Director Prof Lyubomir Penev will shed more light on the first end-to-end publishing solution, providing everything a journal needs in a technologically advanced, highly efficient and user-friendly manner.

ARPHA in a few notes:

The platform

With ARPHA you can choose between two journal publishing workflows: ARPHA-DOC and ARPHA-XML. The former provides document-based submission for the articles in a journal, as well as peer review and publication. The latter makes use of the ARPHA Writing Tool, which takes all processes, including authoring, peer review and post-publication updates, to an online environment, created with collaboration and openness in mind.

Our services

ARPHA takes care of all the steps that go along with academic publishing and its efficient dissemination, so that it provides a long list of perks to make it easier for the scientific community to bring research to light. Website design, online editorial management system, linguistic editing, semantic markup, promotion and (sub-)article usage metrics are only a part of the services the platform has to offer.

The publishing models

ARPHA basically lets users mix-and-match services and features to create the publishing model that’s the best fit for their journals. How do you envision your imprint / web-design look / manuscript input / peer review process / publication output / revenue model? ARPHA understands your journal’s individual needs and works around them.

Find out more about ARPHA at our talk, or come and meet us at our stand in HotSpot Professional and Scientific Information (Hall 4.2, M90).

Mayflies of Turkey: Two new records for the country species and an annotated catalogue

Mayflies (the insect order Ephemeroptera) are a fascinating group, which represents the oldest winged insects, estimated to have been existing on the Earth since the lower Carboniferous, or, approximately for 350 million years. They are characterized by exclusively aquatic larvae, a unique fully winged subimaginal stage (the stage right before the young mayfly transforms into a sexually mature adult) and, typically, rather short life as an adult.

While identification has generally been considered difficult, and good research collections are to be found in relatively few specialised institutions, three biologists from Turkey and Austria have recently concluded a review of the Turkish mayfly fauna, in which they also add two species newly recorded from the country.

They also list 157 mayfly taxa representing 33 genera and 14 families, including 24 species considered endemic to Anatolia. With their annotated overview of the present state of knowledge concerning mayflies in Turkey, the authors aim to facilitate future research.

Synthesis of all previous records of mayflies from Turkey together with new records, a map of provinces and pertinent literature, are all included in the latest paper published by scientists Dr Ali Salur, Hitit University, Çorum, Turkey, Dr Mustafa Cemal Darilmaz, Aksaray University, Aksaray, Turkey, and Dr Ernst Bauernfeind, Natural History Museum Vienna, Austria, in the open access ZooKeys.

The data in the review are based on a detailed study of literature on Ephemeroptera in Turkey as well as on hitherto unpublished material housed in the Natural History Museum Vienna. Unpublished theses have not been considered. By 2015, there have been well over 70 scientific papers and books published on Ephemeroptera in Turkey from both Turkish and foreign researchers.

Distribution of species-group taxa in Turkey have been listed and referenced according to publication dates. National distribution records (without specific data at least on province level) have been listed under ‘Turkey’. Type locality of species were only provided if the taxon had originally been based on material from Turkey. Remarks on different taxonomic opinions and nomenclature have been added under ‘Comment’ whenever appropriate.

Websites http://www.faunaturkey.com and http://www.faunaturkey.org (launched in 2013) are meant to contribute more information on research about the fauna of Turkey. The data provided in the present study will also be added to the websites following publication.

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Original source:

Salur A, Darilmaz MC, Bauernfeind E (2016) An annotated catalogue of the mayfly fauna of Turkey (Insecta, Ephemeroptera). ZooKeys 620: 67-118. doi: 10.3897/zookeys.620.9405

As Good as Gold: Publishing platform ARPHA is proud supporter of Peer Review Week 2016 and Gold star sponsor of Publons’ Sentinels of Science

In the heat of this year’s Peer Review Week, themed “Recognition for Review”, we would like to express how and why we are so proud to be part of it and Publons’ initiative Sentinels of Science, meant to recognize the true guardians of quality science, or in other words, the peer reviewers.

Being a high-tech and modern publishing solution, developed by Pensoft with the mindset that to adapt to the future, means to innovate, ARPHA itself was set to take the quite stagnant current peer review practice forward from day one.

This is why and how we provide a range of peer review options for every author submitting their work to any journal, published on the ARPHA platform. For example, here are the four-stages of the open peer review process operating in our flagship Research Ideas and Outcomes (RIO) Journal:

Author-organised, pre-submission review, available to all journals that make use of our ARPHA Writing Tool, which is our way to take the common get-a-friend-to-proofread-your-work practice to a whole new, transparent and technologically facilitated level. The review happens in real time with the author and the reviewers being able to work together in the ARPHA online environment. It is not mandatory, but we encourage it strongly. All pre-submission reviews provided on authors’ request in RIO can be published along with the article, bearing DOI and citation details.
Pre-submission technical and editorial check is another benefit, provided by the journal’s editorial office to those who are using the ARPHA Writing Tool. If necessary, it can take up several rounds, until the manuscript is improved to the level appropriate for direct submission to the journal.
The community-sourced, post-publication, open peer review is the next review stage provided to all articles published in RIO and all other ARPHA journals.
In addition, RIO also provides journal-organised, post-publication open peer review upon author’s request. In all other ARPHA journals this review stage happens mandatory before publication.

To facilitate peer review in any journal published on the platform, ARPHA consolidates every review automatically into a single online file, which makes it possible for reviewers to comment in real time, even during the authoring process. Once posted, the whole peer review history is archived along with the associated files.

To recognize peer review even further, ARPHA registers automatically each of our peer reviewers, along with their work, on Publons, thanks to the integration of all Pensoft journals with the platform, created to credit reviewers and their contributions.

With this vision of peer review, we simply could not stay clear of the aspiring Sentinels of Science initiative, started by Publons. It only made sense for us to step in, which logically led to the ARPHA logo appearing in the Gold star sponsors list.

On Friday, 23rd September, Publons will announce the recipients of the inaugural Sentinels of Science Award – the top reviewers and editors for the past year. So, tune in this Friday at 4:00 P.M. (BST) and do not forget to join the Twitter conversation via hashtags #PeerRevWk16 and #RecognizeReview.

Greek heroic deity Prometheus now has a namesake in a new tiny rain frog from Ecuador

Prometheus, the mythological Greek heroic deity, has been given a namesake in a new species of tiny rain frog, discovered in southwestern Ecuador. The name was chosen by the international team of scientists, led by Dr Paul Szekely, Ovidius University, Constanta, Romania, in acknowledgement of the Prometeo program, funded by the Ecuadorian government.

The description of this new species (Pristimantis prometeii) is the result of the cooperation between three Romanian Prometeo investigators affiliated with the Universidad Tecnica Particular de Loja and Universidad Nacional de Loja, and two Ecuadorian specialists from Pontificia Universidad Catolica del Ecuador. The full study is available from the open access ZooKeys.

During the day, frogs of the new species were found hiding in flowering plants, while at night — perching on leaves at low heights in well preserved cloud forests. They grow to 2-3 cm with the females being larger than the males.

The newly described species is part of a group of frogs called Terrarana (meaning ‘Land or terrestrial frogs’). This is a lineage of frogs that has evolved directly developing eggs, which are deposited in terrestrial habitats. Unlike other frogs, these ones do not have an aquatic tadpole stage and the embryos develop directly into froglets on land.

The newly described species is only known from Reserva Biologica Buenaventura, southwestern Ecuador, at elevations between 878 and 1082 m. This reserve is privately owned by the Jocotoco Conservation Foundation. The reserve has at least another four endemic species of amphibians, reptiles, and birds.

With more than 470 species, the directly developing rain frogs of the genus Pristimantiscontinue to surprise everyone.

“While new species are described every year, there are over a hundred discovered over the last decade only,” remind the authors.

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Original source:

Szekely P, Cogalniceanu D, Szekely D, Paez N, Ron SR (2016) A new species of Pristimantis from southern Ecuador (Anura, Craugastoridae). ZooKeys 606: 77-97. doi: 10.3897/zookeys.606.9121

A new framework for inferring community assembly processes in ecology

One of the most fundamental goals in ecology – determining the community assembly processes that have structured local communities – has been increasingly studied through the analysis of functional and phylogenetic diversity. Addressing growing evidence that inferences from these measures are often ambiguous to interpret, a new study published in the open access journal One Ecosystem, presents a conceptual framework integrating three approaches which reduces the likelihood of drawing incorrect conclusions.

A long-standing goal in ecology has been to document patterns in natural community structure and connect these patterns to underlying processes such as environmental filtering, disturbance, competition, and predation, collectively known as community assembly processes.

When studying these complex interactions, ecologists have been increasingly leaning towards inferring community assembly processes by calculating communities’ functional and phylogenetic diversity. Unlike diversity measures based solely on species’ identities, functional and phylogenetic diversity can describe important differences between species, ranging across ways in which they use resources, respond to, and influence the local environment.

“Functional and phylogenetic diversity can indeed be useful when trying to infer whether a particular community assembly process, such as interspecific competition or environmental filtering, has been most important for structuring a community or set of communities,” explain the authors of the study.

“However, it is also becoming increasingly obvious that care must be taken when using those methods, since inappropriate choice of traits, diversity metrics, or null models can produce ambiguous results and even lead to incorrect conclusions about the processes underlying community assembly.”

To answer this knowledge gaps, a team of ecologists present a conceptual framework which integrates three approaches to reduce the likelihood of drawing incorrect conclusions from analyses of functional and phylogenetic diversity:

testing hypotheses for how diversity measures and ecological processes vary along an environmental gradient;
analysis of both functional and phylogenetic diversity in concert; and
careful selection of traits related to processes of interest for inclusion in functional diversity analyses.

The new study describes the utility of each of these approaches and shows how combining them can strengthen one’s ability to correctly infer community assembly. The framework is presented in the context of identifying the signatures of interspecific competition and environmental filtering, important processes that operate in many systems across different taxa and are most often referred to in the functional and phylogenetic literature.

“In our paper, we provide examples showing how our framework can be used to test general hypotheses such as the Stress-Dominance Hypothesis,” comment the authors. “Our approach can also be applied to other processes besides competition and environmental filtering. This framework has the potential to enhance comparability between studies and allow for testing of alternative hypotheses regarding changes in community assembly processes along gradients.”

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Original Source:

Lopez B, Burgio K, Carlucci M, Palmquist K, Parada A, Weinberger V, Hulbert A (2016) A new framework for inferring community assembly processes using phylogenetic information, relevant traits and environmental gradients. One Ecosystem 1: e9501. doi: 10.3897/oneeco.1.e9501

Schoolkids name a new tree species from Tanzania while fundraising for tropical forests

A University of York conservationist who put out a call for schools to name a new species of tropical tree has found a winner. Following competitive fundraising totalling nearly £4000, Askham Bryan College, UK pipped the Iringa International School, Tanzania to the post. Coming a close third in the competition were 1st Sherburn Rainbows and Brownies, also from the UK.

Kindly agreeing to share the name with the the African school in second place, the winning college has now baptised the new species Polyceratocarpus askhambryan-iringae.

Dr Andy Marshall, Senior Lecturer in York’s Environment Department and Director of Conservation Science at Flamingo Land, set up the Schools for Forests campaign through his University of York CIRCLE research team after discovering a new species of tree in East Africa.

Discovered by chance in a remote mountain forest in Tanzania while studying one of the world’s rarest primates, the kipunji monkey, Dr Marshall sought to name the new 20-metre tall tree by launching a campaign encouraging schools to raise funds for the conservation of tropical forests.

The winning schools will be twinned with a school in Tanzania and will be offered a free educational trip to Flamingo Land. The new tree’s name was formally announced with anopenly accessible paper in the journal PhytoKeys. And for Flamingo Land, the timing of this species description coincides with another landmark for this North Yorkshire theme park and zoo, where 10,000 native UK trees were recently planted in partnership with the Woodland Trust!

P1140528_AndyMarshallAndXUvariodendron_sp_nov_Vikongwa150mSofCamp_ARM2117 — Dr. Andy Marshall, looking for new species in Tanzanian forests.

“Having the opportunity to name a new species is a truly wonderful thing, which will leave a lasting legacy for all those involved in this campaign. We are so happy to have reached out to so many people, including thousands of people in the UK and Africa through social media”, commented Dr Marshall on the positive experiences around the naming campaign.

“We thank everyone who has been involved in this, especially those who raised these valuable funds, which we will use in our ongoing efforts to save and restore tropical forests.”

“The College has been delighted to be involved in the Schools for Forests campaign. Fundraising by our students included foam parties, raffles and even a human sling shot, and we hope this will be the start of an ongoing partnership!” assed Jane Tupper, Deputy Principal for Teaching & Learning from Askham Bryan College.

Money raised through the Schools for Forests scheme will go towards CIRCLE’s work in Tanzania in collaboration with the Udzungwa Forest Project to better protect tropical forests through scientific research, forest management, community education, training and lobbying.

Original Source:

Marshall AR, Couvreur TLP, Summers AL, Deere NJ, Luke WRQ, Ndangalasi HJ, Sparrow S, Johnson DM (2016) A new species in the tree genus Polyceratocarpus (Annonaceae) from the Udzungwa Mountains of Tanzania. PhytoKeys 63: 63-76. doi: 10.3897/phytokeys.63.6262

Additional Information:

CIRCLE is the Centre for Integrated Research, Conservation and Learning, a collaboration between the University of York’s Environment Department and Flamingo Land that aims to protect biodiversity, animal welfare and promote public understanding of the natural world.

Sponsors of the Schools for Forests campaign include United Bank of Carbon, the British Association of Zoos and Aquariums Mammal Working Group and Santander. The Schools for Forests team are also thankful for support from Natalie Bennett, Green Party Leader, and from various collaborators, especially David Johnson from Ohio Wesleyan University.

Poorly known South African mountain endemic appears to be a very valuable keystone species

Mountain ecosystems are valuable providers of key resources including water. These ecosystems comprise diverse species, some of which appear to be especially important to the ecosystem’s functioning. In poorly studied mountain environments in biodiversity-rich countries, these keystone species can often be overlooked and undervalued.

Macowania is a group of yellow daisy shrubs occurring in the alpine-like regions of the Drakensberg and highlands of Ethiopia, Eritrea and Yemen. Doctoral student Joanne Bentley, University of Cape Town, studied the genetic relationships between the various Macowaniaspecies and relatives during her Masters degree studies. Her research led to the first collection of the poorly known species Macowania revoluta (known also as the Amathole Macowania) in about 40 years.

The story of Macowania revoluta is published in the open access journal PhytoKeys.

The Amathole Macowania appears to be an exceptionally important keystone species. This is because it forms one of the dominant members of the valuable mountain wetland communities and, thus, likely plays a very important role in wetland functioning and soil protection.

It appears to be somewhat tolerant of woody alien species and a valuable pioneer species protecting its native co-habitants. Plants like this one buffer more sensitive plants from sudden changes in environment (such as forestry, alien invasion and fire), and provide an opportunity for the ecosystem to ‘bounce back’.

Restricted to the Amathole mountains in the Eastern Cape Province, South Africa, the Amathole Macowania was first collected sometime before 1870 by the pioneer botanist Peter MacOwan, and was well documented until around 1949. After that, except for one record in 1976, the plant quietly disappeared.

“This was the first Macowania species that we found during our fieldtrip across the greater Drakensberg. We had combed several of the localities where it had been collected before; mostly from several decades ago, some from more than a century ago!” says Joanne Bentley. “We became increasingly doubtful about finding the plant, given the heavily transformed plantation landscape.”

“Ready to throw in the towel, we came across a peaty area on the margins of the forest and decided on one last investigation. We were lucky: it was growing prolifically! It was a very special moment.”

As it often happens, exciting discoveries come in bulk. Joanne’s discovery of the plant in July 2010 was followed by another record in October 2010, by the Curator of the Schonland Herbarium, Tony Dold. In 2014 at least three additional localities were recorded along the popular Amathole Hiking Trail by Dr Ralph Clark, Rhodes University. A further record was added in 2015 by Vathi Zikishe, South African National Biodiversity Institute. The verdict: this is a very localised but patchily abundant species, and an ecologically valuable component of the Amathole flora.

Listed as ‘Data Deficient’ in the Threated Plants List for South Africa, this string of modern records of the species also provided the first opportunity to get an idea of its ecology and abundance, as well as the first photographs.

“The practical value of this species in local land restoration projects still needs to be explored, but the opportunities are exciting,” says Dr Clark. “The discovery that this obscure endemic mountain plant is not only abundant, but is, in fact, fulfilling an extremely important ecological role, highlights the value of detailed mountain biodiversity research in southern Africa.”

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Original source

Clark VR, Bentley J, Dold AP, Zikishe V, Barker NP (2016) The rediscovery of the Great Winterberg endemic Lotononis harveyi B.-E.van Wyk after 147 years, and notes on the poorly known Amathole endemic Macowania revoluta Oliv. (southern Great Escarpment, South Africa). PhytoKeys 62: 1-13. doi: 10.3897/phytokeys.62.8348

Surprising exotic flies in the backyard: New gnat species from Museum Koenig’s garden

Little did scientists Kai Heller and Björn Rulik expect to discover a new species in Germany’s Alexander Koenig Museum‘s garden upon placing a malaise trap for testing purposes. Not only did an unknown and strikingly coloured gnat get caught, but it turned out to be a species, which showed to have much more in common with its relatives from New Zealand. Their study is published in the open access Biodiversity Data Journal (BDJ).

While the genus, which the new dark-winged fungus gnat species belongs to, likely originates from the Australasian region, it was so far represented by only three species in Europe. None of them, however, stands out with the contrasting colouration of the presently announced fourth one.

The new gnat, called Ctenosciara alexanderkoenigi after the German museum’s founder, is described based on a single specimen caught in the framework of the German Barcode of Life Project (GBOL). Over three days, the scientists observed the flying insects getting caught in a malaise trap, placed among the predominantly non-native plants in the Alexander Koenig Museum’s garden. This tent-like structure is designed to catch flying insects. Once they fly into its walls, they get funnelled into a collecting bottle.

Upon noticing the beautiful striking colour of the fly, the two specialists were convinced they had just discovered a new to science species. Most of these flies are bright brownish, and the only other orange European dark-winged fungus gnat – almost uniformly orange. In contrast, the new species stands out with a mixture of reddish, black and yellowish-white hues. Based on the DNA-barcode match with New Zealand specimens, the authors concluded that the species must have arrived from the Australasian region in Europe quite recently.

“It is a rare occurrence, that a species from the opposite end of the world is represented by a single specimen only and it is not yet clear, whether Ctenosciara alexanderkoenigi has a permanent population in Germany or if it was only introduced casually with plants or soil,” they explain. “Probably, the species was recently introduced from the Australasian Region. If it was a permanent member of the European fauna, a striking species like this would likely have been found earlier.”

In conclusion, the scientists note that modern technologies such as the high quality photo documentation, established as a standard by the BOLD project, DNA barcodes assigned with BINs, as well as facilitated by speedy publishing, have largely aided taxonomists to build on the biodiversity knowledge.

“We believe that the rapid description of Ctenosciara alexanderkoenigi, coupled with the BDJ reviewing system, might be a robust and ground-breaking way to accelerate and stabilise taxonomy in the future,” they finish their paper.

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Original source:

Heller K & Rulik B (2016) Ctenosciara alexanderkoenigi sp. n. (Diptera: Sciaridae), an exotic invader in Germany? Biodiversity Data Journal 4: e6460. doi: 10.3897/BDJ.4.e6460

Two brand new dung beetle species from montane grazing sites and forests in Mexico

While carrying out a biodiversity study, a Mexican-Italian research team discovered three new dung beetle species in montane forests disturbed by livestock grazing. Mexico has been a mecca for naturalists, and its dung beetle species are among the best known in the world. This is why the discovery of new species there is noteworthy. The present study, published in the open-access journal ZooKeys, describes two of them and highlights the need to further explore the biodiversity of disturbed ecosystems.

Mexico is a country that holds a vast number of creatures and ecosystems. There is in fact a fascinating phenomenon: tropical forests that have close affinities with South America co-occurring with temperate and arid areas shared with North America. Thus, Mexico has been particularly attractive to explorers ever since the 19th century.

A group of animals that has woken up a special interest for studies in Mexico is the so-called ‘dung beetles’. As their name suggests, dung beetles are insects that feed mainly on mammal faeces.

For decades, an international research team, led by Dr Gonzalo Halffter, has studied dung beetles across the world, especially in Mexico. As a consequence, the Mexican species are some of the best-known. However, Dr Halffter and his team are not interested exclusively in dung beetles, but also in evolutive phenomena, the effects of land-use change, ecosystems modification by human activities, and conservation biology. Such concerns seem to be of particular importance now that the terrestrial ecosystems in Mexico have been severely destroyed and disturbed by people.

Livestock is one of the major drivers of biodiversity loss worldwide, which makes the present discovery particularly impressive. With at least 58% of the area of Mexico occupied with livestock farming, dung beetles are essential in cleaning up. While studying their diversity at conserved forests and cattle grazing sites across the mountains of Mexico, the researchers found some new species of dung beetles.

The first to discover these new dung beetles was Victor Moctezuma, a student of Dr Gonzalo’s at the Instituto de Ecología of Mexico.

“I was carrying out sampling for my Masters Degree studies, but I had no idea that new dung beetles could be found in a forest that is disturbed by human activities, such as livestock grazing and land-use change,” recalls Moctezuma. “So I was really surprised when I discovered three dung beetle species.” One of these species has already been published.

Apart from the two new dung beetles, formally called Onthophagus clavijeroi and Onthophagus martinpierai, the present paper also provides theories about the current distributions of these insects across the Mexican mountains and their putative evolutive relationships. As a whole, the study highlights the importance of disturbed forest for species discovery and conservation.

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Original source:

Moctezuma V, Rossini M, Zunino M, Halffter G (2016) A contribution to the knowledge of the mountain entomofauna of Mexico with a description of two new species of Onthophagus latreille, 1802 (Coleoptera, Scarabaeidae, Scarabaeinae). ZooKeys 572: 23-50. doi:10.3897/zookeys.572.6763

Zorro, the new Latin American fish species, takes off the mask to show its true identity

Unidentified since its discovery in 2007, a large fish species from Amazonia has failed to give out enough information about itself, leaving only insufficient hints about its genus. Nevertheless, three scientists have now recovered the missing pieces to puzzle out its mysterious identity. In their study, published in the open-access journal ZooKeys, they describe the fish as a new species and name it after the fictional secretive Latin American character Zorro.

The new fish, called Myloplus zorroi, is commonly known among the Brazilians as ‘pacu’ and is a relative to the piranha. The research team, led by Marcelo C. Andrade, Universidade Federal do Para, Brazil, recognised in a fish, collected by sport fishermen from Rio Madeira basin, Brazil, a previously found, yet undescribed species. Following their analysis, it turned out that its discoverers had assumed an incorrect genus for it.

Among the distinctive features of the new fish, which helped its rightful placement, are its characteristic teeth, specialised to crush seeds.

The new pacu species is quite large, growing up to 47,5 cm. It dwells in moderately to rapidly flowing clear rivers, running over rocky or sandy bottoms, and ranging from about 2 to 8 metres in depth. Its basis colour is reddish silver with darker markings running along the upper side of the body. The head is dark and the belly – pale yellow.

Curiously enough, although the name of the new fish is chosen as a tribute to Mauricio Camargo-Zorro, a researcher at the Instituto Federal de Educacao, Ciencia e Tecnologia, in recognition of his invaluable contribution to the fish fauna inventory from the Marmelos Conservation Area, zorroi is also a playful reference to the Latin American fictional character Don Diego de la Vega and his secret identity hidden behind the nickname of Zorro.

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Original source:

Andrade MC, Jegu M, Giarrizzo T (2016) A new large species of Myloplus (Characiformes, Serrasalmidae) from the Rio Madeira basin, Brazil. ZooKeys 571: 153-167. doi: http://doi.org/10.3897/zookeys.571.5983.